animations body is not compatible with skeleton

[MarkoGrbec]

Tested versions

not reproducible in Godot 4.6 Dev-2
reproducible in Godot 4.6 Dev-3
reproducible in Godot 4.6 Dev-4

System information

Godot v4.6.dev4 - Manjaro Linux #1 SMP PREEMPT_DYNAMIC Fri, 19 Sep 2025 16:09:36 +0000 on X11 - X11 display driver, Multi-window, 1 monitor - Vulkan (Mobile) - integrated Intel(R) HD Graphics 4600 (HSW GT2) - Intel(R) Core(TM) i7-4700HQ CPU @ 2.40GHz (8 threads) - 15.49 GiB memory

Issue description

animations are still compatible with skeleton but skeleton isn't compatible with mesh any longer.

Steps to reproduce

you must run 4.6 Dev2 to be working anmations with mesh

if you run 4.6 Dev 3 or 4 mesh doesn't follow any longer.

Minimal reproduction project (MRP)

testing_2025-09-23_16-50-51.zip

[AThousandShips]

This seems like a regression from:

• Remove default skeleton path in MeshInstance3D #112267

Which should be fixed either of:

• Add MeshInstance3D upgrade code #112607
• Revert "Remove default skeleton path in MeshInstance3D" #112605

[patwork]

[akien-mga]

You can fix the issue by reimporting the glb in 4.6-dev3 or later (either manually, or delete .godot to reimport everything).

#112267 changed way skeleton paths are stored in this specific scenario where the MeshInstance3D is under a parent Skeleton3D, which is the case in your glTF file. But this information is already part of the glTF so reimporting it fixes the issue.

[MarkoGrbec]

ok thanks it works.

this really should be somewhere in the docs.
re importing only mesh worked fine.

is it possible to get a error in console? to re import. kinda hard I think don't know.

[patwork]

There is no official migration guide yet, but remember that you are using a dev snapshot, so many things may still change.

https://docs.godotengine.org/en/stable/tutorials/migrating/index.html